| #1 | Harvard University(Cambridge, MA) | United States |
| #2 | Massachusetts Institute of Technology(Cambridge, MA) | United States |
| #3 | University of Washington(Seattle, WA) | United States |
| #4 | University of Oxford(Oxford) | United Kingdom |
| #5 | University of California–San Francisco(San Francisco, CA) | United States |
| #6 | Duke University(Durham, NC) | United States |
| #7 | Rockefeller University(New York, NY) | United States |
| #8 | University of Wisconsin–Madison(Madison, WI) | United States |
| #9 | Washington University in St. Louis(St. Louis, MO) | United States |
| #10 | University of Pennsylvania(Philadelphia, PA) | United States |
| #11 | Johns Hopkins University(Baltimore, MD) | United States |
| #11 | Stanford University(Stanford, CA) | United States |
| #13 | University of California–Berkeley(Berkeley, CA) | United States |
| #14 | University of North Carolina–Chapel Hill(Chapel Hill, NC) | United States |
| #15 | Imperial College London(London) | United Kingdom |
| #16 | Cornell University(Ithaca, NY) | United States |
| #16 | University of California–San Diego(La Jolla, CA) | United States |
| #18 | Emory University(Atlanta, GA) | United States |
| #19 | University of Tokyo(Bunkyo-ku, Tokyo) | Japan |
| #20 | University of Cambridge(Cambridge) | United Kingdom |
| #21 | Yale University(New Haven, CT) | United States |
| #22 | Columbia University(New York, NY) | United States |
| #23 | University of Edinburgh(Edinburgh, Scotland) | United Kingdom |
| #24 | University of California–Los Angeles(Los Angeles, CA) | United States |
| #25 | Mount Sinai School of Medicine(New York, NY) | United States |
| #26 | Erasmus University Rotterdam(Rotterdam) | Netherlands |
| #27 | University of Hong Kong(Pok Fu Lam, Hong Kong) | Hong Kong |
| #28 | University of Glasgow(Glasgow, Scotland) | United Kingdom |
| #28 | University of Queensland Australia(Brisbane, Queensland) | Australia |
| #30 | McGill University(Montréal, Québec) | Canada |
| #31 | University of Toronto(Toronto, Ontario) | Canada |
| #32 | University of Maryland–Baltimore(Baltimore, MD) | United States |
| #33 | University of Alabama–Birmingham(Birmingham, AL) | United States |
| #33 | University of Melbourne(Parkville, Victoria) | Australia |
| #35 | London School of Hygiene & Tropical Medicine(London) | United Kingdom |
| #36 | Karolinska Institute(Stockholm) | Sweden |
| #37 | University of Copenhagen(Copenhagen) | Denmark |
| #37 | University of Montpellier(Montpellier) | France |
| #39 | University of Aix-Marseille(Marseille) | France |
| #39 | University of Michigan–Ann Arbor(Ann Arbor, MI) | United States |
| #41 | New York University(New York, NY) | United States |
| #41 | Swiss Federal Institute of Technology Zurich(Zurich) | Switzerland |
| #43 | Pennsylvania State University–University Park(University Park, PA) | United States |
| #44 | University of Chicago(Chicago, IL) | United States |
| #45 | University College London(London) | United Kingdom |
| #45 | University of Pittsburgh(Pittsburgh, PA) | United States |
| #47 | University of Georgia(Athens, GA) | United States |
| #47 | University of São Paulo(São Paulo) | Brazil |
| #49 | University of California–Davis(Davis, CA) | United States |
| #50 | Seoul National University(Seoul) | South Korea |
| #50 | University of Munich(Munich) | Germany |
| #52 | University of Texas Medical Branch–Galveston(Galveston, TX) | United States |
| #53 | University of Helsinki(Helsinki) | Finland |
| #54 | Heidelberg University(Heidelberg) | Germany |
| #55 | University of Liverpool(Liverpool) | United Kingdom |
| #56 | Osaka University(Suita, Osaka) | Japan |
| #56 | University of Amsterdam(Amsterdam) | Netherlands |
| #58 | Catholic University of Leuven(Leuven) | Belgium |
| #58 | Vanderbilt University(Nashville, TN) | United States |
| #60 | Ghent University(Gent) | Belgium |
| #61 | National University of Singapore(Singapore) | Singapore |
| #61 | University of Minnesota–Twin Cities(Minneapolis, MN) | United States |
| #63 | Wageningen University and Research Center(Wageningen) | Netherlands |
| #64 | Leiden University(Leiden) | Netherlands |
| #64 | Princeton University(Princeton, NJ) | United States |
| #66 | University of Southern California(Los Angeles, CA) | United States |
| #67 | University of Colorado–Boulder(Boulder, CO) | United States |
| #68 | University of Florida(Gainesville, FL) | United States |
| #69 | Ohio State University–Columbus(Columbus, OH) | United States |
| #70 | Kyoto University(Sakyo-ku, Kyoto, Kyoto) | Japan |
| #70 | Monash University(Clayton, Victoria) | Australia |
| #72 | University of Vienna(Vienna) | Austria |
| #73 | University of Zurich(Zürich) | Switzerland |
| #74 | Pierre and Marie Curie University(Paris) | France |
| #74 | Utrecht University(Utrecht) | Netherlands |
| #76 | Paris Descartes University-Paris V(Paris) | France |
| #76 | University of Würzburg(Würzburg) | Germany |
| #78 | University of Freiburg(Freiburg) | Germany |
| #79 | University of British Columbia(Vancouver, British Columbia) | Canada |
| #79 | University of Strasbourg(Strasbourg) | France |
| #79 | Yeshiva University(New York, NY) | United States |
| #82 | Hokkaido University(Kita-ku, Sapporo, Hokkaido) | Japan |
| #83 | University of California–Irvine(Irvine, CA) | United States |
| #84 | Rutgers, The State University of New Jersey–New Brunswick(Piscataway Township, NJ) | United States |
| #85 | University of New South Wales(Kensington, New South Wales) | Australia |
| #85 | University of Sydney(Sydney, New South Wales) | Australia |
| #87 | Mahidol University(Salaya, Nakhon Pathom) | Thailand |
| #88 | University of Cape Town(Cape Town) | South Africa |
| #88 | University of Massachusetts Worcester(Worcester, MA) | United States |
| #90 | University of Aberdeen(Aberdeen, Scotland) | United Kingdom |
| #91 | University of Lausanne(Lausanne) | Switzerland |
| #92 | Radboud University Nijmegen(Nijmegen) | Netherlands |
| #93 | Case Western Reserve University(Cleveland, OH) | United States |
| #94 | Purdue University–West Lafayette(West Lafayette, IN) | United States |
| #94 | University of Geneva(Geneva) | Switzerland |
| #96 | Freie Universität Berlin(Berlin) | Germany |
| #97 | Hannover Medical School(Hannover) | Germany |
| #98 | University of Basel(Basel) | Switzerland |
| #99 | Hebrew University of Jerusalem(Jerusalem) | Israel |
| #100 | Northwestern University(Evanston, IL) | United States |
